The upper molar cusps is actually seemingly bunoselenomorph

They have a short and you may straight postprotocrista labially dependent. The newest preprotocrista, seemingly straight, satisfies a properly-establish parastyle. The metaconule is a bit reduced which is larger to your M3. The new premetaconulecrista therefore the fossae are pretty straight forward in the place of neocrista. Brand new rear fossa was angled, especially toward M3. Off upright towards the M1, the fresh new postmetaconulecrista gets sigmoidal into the M3, bypassing the fresh metacone. The brand new labial cusps come in a similar axis having relatively upright cristae. This new paracone labial rib are salient, however, versus anterior groove. There is no metacone labial rib. The brand new para poder-, meso-, and metastyles was outstanding. The fresh new lingual cingulum is quite weakened so you’re able to absent for instance the entostyle.

The latest submitted structure of your next straight down premolar is normal to own derived stalk pecoran and you can top pecoran for the which have a well-arranged mesolingual conid and you will posterior cristid , . Moreover, the clear presence of a tooth retailer towards p1 excludes an enthusiastic attribution to help you Prodremotherium otherwise Mosaicomeryx . Even when the white teeth aren’t as the bunoselenodont as in Amphitragulus elegans, the fresh new referred specimens have very primitive have, for example a very quicker metastylid, a low crown, a little postentocristid, and although laterally compacted, bulged lingual cuspids to your lower molars. All these qualities exclude an affiliation to help you sometimes Dremotherium or Bedenomeryx , . Furthermore, the new very small-size off “Amphitragulus” quercyi try a pinpointing function into the newest Oligocene ruminants away from European countries , , , .

Dremotherium cf. guthi regarding Gaimersheim1: step 1 left maxillary limbs with the enamel line M1-M3 (SNSB-BSPG 1952II4855). from Mailhat: dos kept P4-M2 (MNHN LIM154). Babameryx engesseri off Gaimersheim1: 3 leftover fragmentary mandible that have m2 (SNSB-BSPG 1952II4818), (a) lingual, (b) labial, and you can (c) occlusal views. The scale club was 2 cm.

Holotype called specimens recommendations

Holotype. Jehenne (1987) enjoys appointed the fresh new specimen IPHEP LM.1969.MA.forty because the holotype. This really is a partial right maxillary bone which have P3-M3 (, Fig. 2), of Los angeles Milloque (MP29, France).

Referred specimens. Mailhat MNHN LIM154 remaining P4-M2 (that it works, Fig. 6.2); Gaimersheim1 SNSB-BSPG 1952II4855 kept maxillary with M1-M3 (it really works, Fig. 6.1); SNSB-BSPG 1952II4870 correct m2.


Lower permanent dentition. The low molar is quite defectively preserved. However, this big ruminant and also a totally selenomorph trigonid. The newest cristid is actually elongated as well as the cuspids try highest towards the enamel a little pleated. The fresh metastylid was ripped.

Upper long lasting dentition. The fresh new P4 is actually defectively kept. All the lingual region try destroyed. The brand new anterolabial cone features a faltering labial rib. The prior stylid try busted. New rear you’re brief however, establish. Zero neocrista obstructs the fossa. The fresh quadratic higher molars during the occlusal take a look at incur a proper-set-up metaconule some smaller than this new protocone. The newest cusps have become selenomorph. Brand new preprotocrista together with postmetaconulecrista was labially oriented and you can a little upright. The fresh postprotocrista is long, highly rounded, and you can labially built. It may be totally isolated (SNSB-BSPG 1952II4855) otherwise bonded (MNHN LIM154) towards premetaconulecrista. The second ends up during the axis of one’s mesostyle. The posterior basin can provides a little metaconule fold with the M2, big on the M3. The paracone, having an effective paracone rib, additionally the metacone, labially apartment and you can concave, try lined up. This new mesostyle and parastyle are developed. It mode rounded brief columns. This new mesostyle try huge regarding M1 so you can M3. Brand new entostyle try poor otherwise missing. Discover none a beneficial lingual cingulum nearby the new protocone neither a great metastyle creating a properly-developed cuspid. Yet not, brief anterior and rear cingula exists.


This ruminant is fairly cutting-edge in being well-selenodont and achieving no lingual cingula from inside the higher molars, usually contained in Lophiomeryx, Gelocus, Iberomeryx, Mosaicomeryx, and Babameryx , , , [61–63], . In comparison to Bachitherium, so it varieties does not have a keen prior groove on very big labial paracone rib , , . The newest examined fossils much more much like the cutting-edge base Pecora and this new primitive crown Pecora (such as for instance Amphitragulus, Dremotherium, Bedenomeryx). The new tooth was somewhat pleated, while it’s generally simple within the Prodremotherium. Additionally, how big is such white teeth is significantly bigger than from inside the Prodremotherium from the dated Quercy series and you can Pech Desse . Tooth be much more selenomorph than in “Amphitragulus” and Bedenomeryx , , as well as the parastyle is not that anteriorly projected than just noticed for the “Amphitragulus” feningrei , , . Taking a look at the neodiagnosis advised of the Mennecart , this type of fossils were all exhibiting top features of Dremotherium. New currently defined types into the European countries try D. feignouxi (types of varieties) and D. guthi , , . The latter was slightly smaller than the type species , much like the fossils explained here. D. feignouxi just occurred in Agenian (MN1-MN2; ), while D. guthi resided in the current Oligocene (MP28-MP30; ). Although not, within the absence of extra better-maintained procedure, a decisive assignation towards kinds isn’t feasible.